List of publications published as part of the Bhutan Biodiversity Project:
Dorji, C., P. Loday & O. Vorst (2017). A field guide to the common Lady Beetles of Bhutan. National Biodiversity Centre (NBC), Ministry of Agriculture & Forest, Serbithang, Thimphu, Bhutan.
Dorji, C., P. Loday & O. Vorst (2019). A preliminary checklist of the Coccinellidae of Bhutan (Insecta: Coleoptera). Zootaxa 4712 (5): 497-530.
Dorji, K, S. Tashi, J.C. Biesmeijer, N. Leclercq, J. Nicolas, L. Marshall (2022). Pollinators and crops in Bhutan: insect abundance improves fruit quality in Himalayan apple orchards. Journal of Pollination Ecology 31: 39-52.
Dorji, P., T. Gyeltshen, W. Klein & T. Nidup (2017). New records of social wasps (Hymenoptera: Vespinae: Vespa and Provespa) from Bhutan. Journal of Threatened Taxa 9(4): 10102-10108.
Dorji, P., T. Nidup & W. Klein (2017). A field guide to the common bees & wasp of Bhutan. National Biodiversity Centre (NBC), Ministry of Agriculture & Forest, Serbithang, Thimphu, Bhutan.
Dorji, P., W. Klein & T. Nidup (2016). New record of Carpenter bees (Hymenoptera: Apidae: Xylocopinae) from Bhutan. Journal on New Biological Reports 5: 106-113.
Dorji, P., W. Klein & T. Nidup (2016). Study of Paper wasps (Hymenoptera: Vespidae: Polistinae) of Bhutan. Bio Bulletin 2: 128-136.
Dorji, P., W. Klein & T. Nidup (2017). Taxonomic study of social vespid wasps (Hymenoptera: Vespidae: Vespinae & Polistinae) in Bhutan. Journal of Insect Biodiversity and Systematics, 3(2): 91-104.
Dorji, P., T. Nidup, W. Klein, C. Dorji, A. Daglio & C. Gyeltshen (2022). An annotated checklist of the Vespidae (Hymenoptera: Vespoidea) of Bhutan with new records. Zootaxa 10.11646/zootaxa.5150.3.1
Gerecke, R. & H. Smit (2022). Water mites of the genus Lebertia Neuman, 1880 from the eastern Himalayas (Acari: Hydrachnidia: Lebertiidae). Acarologia, 62(2), 302-316. https://zoobank.org/52668A17-CAAF-437A-864C-A2883EF87409.
Gielis C. & K. Wangdi (2019). The Pterophoridae of Bhutan (Lepidoptera), with the description of a new species. Tijdschrift voor Entomologie 161: 79-109.
Gielis, C. & K. Wangdi (2017). Common Moths of Bhutan. National Biodiversity Centre (NBC), Ministry of Agriculture & Forest, Serbithang, Thimphu, Bhutan.
Gielis, K., M. Franssen, F. Groenen & K. Wangdi (2022). Moths of Bhutan. Gielis & Klein publishers. 420 pp.
Gittenberger E, S. Sherub, B. Stelbrink (2017). Erhaia Davis & Kuo (Gastropoda, Rissooidea, Amnicolidae) also in Bhutan. ZooKeys 679: 21-28.
Gittenberger, E., C. Gyeltshen& S. Sherub (2021). Conspicuously sculptured small shells from Bhutan: Philalanka bhutana sp. nov. and Sculpteuconulus obliquistriatus gen. et sp. nov. Archiv für Molluskenkunde 150: 5-11.
Gittenberger, E., C. Gyeltshen, K. Tobgay & S. Sherub (2022). The genera Dioryx and Cycloryx (Gastropoda, Caenogastropoda, Alycaeidae) in Bhutan, with a description of four new species. Basteria 86 (2) in print.
Gittenberger, E., C. Gyeltshen, K. Tobgay & S. Sherub, S. (2022). High diversity of Endothyrella (Gastropoda, Pulmonata, Plectopylidae) in Bhutan, with a description of four new species. Basteria 86 (2) in print.
Gittenberger, E., C. Gyeltshen, P. Leda, T. Zangpo, R. Dekker van Klinken (2018). The first record of the cosmopolitan slug Deroceras laeve (O.F. Müller, 1774) (Gastropodae, Pulmonata, Agriolimacidae) in Bhutan. Folia Malacologica 26 (2): 1-5.
Gittenberger, E., Leda, P., Gyeltshen, C. & S. Sherub (2018). Distributional patterns of molluscan taxa in Bhutan (Mollusca). Biodiversität und Naturausstattung im Himalaya VI. Verein der Freunde & Förderer des Naturkundemuseums Erfurt eV, Erfurt: 143-151.
Gittenberger, E., P. Leda & S. Sherub (2017). Das malakologisch unbekannte Bhutan erstmalig erschlossen. Mitteilungen der Deutschen Malakozoologischen Gesellschaft 96: 48.
Gittenberger, E., P. Leda, S. Sherub & C. Gyeltshen (2019). The subfamily Phaedusinae in Bhutan (Gastropoda, Pulmonata, Clausiliidae). Basteria 83 (4-6): 133-144.
Gittenberger, E., P. Leda, J. Wangchuk, C. Gyeltshen & B. Brink (2020). The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17.
Gittenberger, E., P. Leda, A.J. Winter de & A. Jochum (2017). First record of Carychium in Bhutan (Gastropoda, Ellobioidea). Basteria 82 (4-6): 107-110.
Gittenberger, E., P. Leda & S. Sherub (2013). Bhutanese snails, the smallest one: Truncatellina bhutanensis spec. nov. (Gastropoda, Pulmonata, Vertiginidae). Basteria 77: 29-32.
Gittenberger, E., P. Leda & S. Sherub (2017). Gastropods in Bhutan, the genus Rahula (Pulmonata: Helicarionidae). Journal of Conchology 42: 413-418.
Gittenberger, E., P. Leda, C. Gyeltshen, S. Sherub & S. Dema (2017). Field Guide to the Common Molluscs of Bhutan. National Biodiversity Centre (NBC), Ministry of Agriculture & Forest, Serbithang, Thimphu, Bhutan.
Gittenberger, E., P. Leda, K. Wangdi & S. Sherub (2017). Bhutan freshwater gastropods and trematodes, with a warning. Biodiversity Journal 8: 895-906.
Gittenberger, E., P. Leda, S. Sherub & B. Páll-Gergely (2018). Endothyrella Zilch, 1960 in Bhutan (Gastropoda: Pulmonata: Plectopylidae), with a description of three new species (2018). Archiv fur Molluskenkunde 147: 203-213.
Gittenberger, E., P. Leda, S. Sherub & C. Gyeltshen (2021). Rahula revisted (Pulmonata: Euconulidae), with data for Bhutan, India (Assam), Laos, Vietnam and Indonesia, including two new species. Journal of Conchology 44: 11-19.
Gittenberger, E., P. Leda, S. Sherub & C. Gyeltshen (2021). The families Streptaxidae and Diapheridae (Gastropoda, Pulmonata) in Bhutan, with notes on some species occurring in Nepal and North-east India. Basteria 85: 73-81.
Gittenberger, E., T. Tobgay, C. Gyeltshen & H. Nesemann (2021). Two high-altitude species of molluscs, new for Bhutan (Bivalvia, Sphaeriidae - Gastropoda, Lymnaeidae). Basteria 85: 13-17.
Gittenberger, E., U. Tenzin & S. Sherub (2017). Additional records of Rahula species (Pulmonata, Helicarionidae) in Bhutan. Basteria 81 (4-6): 119-122
Gittenberger, E., C. Gyeltshen, P. Leda & S. Sherub (2021). The superfamilies Pupilloidea and Enoidea (Gastropoda, Eupulmonata) in Bhutan. Folia Malacologica 29(2): 69-90.
Gittenberger E, C. Gyeltshen, B. Stelbrink (2022). The genus Erhaia (Gastropoda, Truncatelloidea, Amnicolidae), with a new species from Bhutan. Zookeys 1085: 1-9.
Groenen, F. & K. Wangdi (2019). The Tortricidae (Lepidoptera) of Bhutan. Zootaxa 4658: 526-540.
Gurung, M.M., V.J. Kalkman, G.S. Bhandari & A. Dhimal (2021). Nine new species of dragonfly and damselfly for Bhutan (Insect: Odonata) with a note on Calicnemia mortoni. Agrion 25(1): 22-28.
Gurung, M.M., C. Dorji, A.M. Sinchuri, S.K. Rai, K.C. Dendup & V.J. Kalkman (2022). New records of odonates from Trongsa and Zhemgang, central Bhutan with a checklist of Jigme Singye Wangchuck National Park. Journal of Threatened Taxa 14(9): 21836-21844.
Gurung, M.M., C. Dorji, D.B. Gurung & H. Smit (2022). Checklist of water mites (Acari: Hydrachnidia) of the Himalayan and Tien Shan Mountains. Ecologica Montenegrin 57: 8-23.
Gurung, M.M., C. Dorji, D.B. Gurung & H. Smit (2022, in print). Environmental factors affecting the water mites (Acari: Hydrachnidia) assemblage in streams, Mangde Chhu basin, Central Bhutan. Journal of Threatened Taxa.
Gyeltshen, T. & V.J. Kalkman (2017). Odonata records from western Bhutan, with six new records and a note on the synonymy of Himalagrion with Coenagrion. Notulae odonatologicae 8(9): 341-353.
Gyeltshen, T. (2017). A survey of Odonata from eastern Bhutan, with nine new national records. Notulae odonatologicae 89: 319-374.
Gyeltshen, T., T. Nidup, P. Dorji & V.J. Kalkman (2016). A bibliography and checklist of the dragonflies and damselflies of Bhutan. Journal of Threatened Taxa 9: 9743-9747. PDF
Gyeltshen, T., V.J. Kalkman & A.G. Orr (2017). A field guide to the common Dragonflies & Damselflies of Bhutan. National Biodiversity Centre (NBC), Ministry of Agriculture & Forest, Serbithang, Thimphu, Bhutan.
Gyeltshen, T., V.J. Kalkman & A.G. Orr (2017). Honouring His Royal Highness the Crown Prince of Bhutan: Megalestes gyalsey (Odonata: Synlestidae). Zootaxa 4244: 588-594.
Gyeltshen, T. (2020). Eight new additional records of Odonata from Bhutan. Agrion. 24(2), 97-103.
Groenen, F. & K. Wangdi (2019). The Tortricidae (Lepidoptera) of Bhutan. Zootaxa 4658 No.3 pp.526-540.
Kalkman, V.J. & T. Gyeltshen (2016). Records of dragonflies from Western Bhutan collected in October 2015. IDF-reports 94: 1-15.
Kalkman, V.J., R. Babu, M. Bedjanic, K. Conniff, T. Gyeltshen, M.K. Khan, K.A. Subramanian, A. Zia & A.G. Orr (2020). Checklist of the dragonflies and damselflies (Insecta: Odonata) of Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka. Zootaxa 4849(1): 1-84.
Kalkman, V.J., M.M. Gurung & H. Zhang (2022). On the Asian species of Perissogomphus Laidlaw, 1922, and Ophiogomphus Selys, 1854 (Odonata: Gomphidae). Notulae odonatologicae 9 (9), 389-397.
Nidup, T. & P. Dorji (2016). The Honey Bees (Hymenoptera: Apidae) of Bhutan with a key to the Apis species. Bio Bulletin 2: 1-7.
Nidup, T., T. Gyeltshen, P. Girish Kumar, W. Klein & P. Dorji (2016). New records of petiolate potter wasps (Hymenoptera: Vespidae: Eumeninae) from Bhutan. Journal of Threatened Taxa 8 (12): 9467-9472.
Nidup, T., W. Klein & P. Dorji (2016). New records of hover wasps (Hymenoptera: Vespidae: Stenogastrinae) from Bhutan. Journal of Threatened Taxa 9 (7): 10487-10489.
Nidup, T., W. Klein, P.G. Kumar & P. Dorji (2017). New record of scoliid wasps (Hymenoptera: Scoliidae: Scoliinae) from Bhutan. Halteres 8: 9-19.
Nidup, T., W. Klein, P.G. Kumar & P. Dorji (2018). Additional reports of solitary potter wasps (Hymenoptera: Vespidae: Eumeninae) in Bhutan. Journal of Threatened Taxa 10(5): 11686-11696. DOI: 10.11609/jott.3888.10.5.11686-11696
Pešić, V., H. Smit & M.M. Gurung (2022). Torrenticolid water mites of Bhutan. Genera Torrenticola Piersig, 1896 and Neoatractides Lundblad 1941 (Acari; Hydrachnidia: Torrenticolidae). Acarologia, 62(3), 821-860. https://doi.org/10.24349/xn0u-5px2.
Pešić, V., H. Smit & M.M. Gurung (2022). Neumania bhutana sp. nov. a new water mite from Bhutan (Acari, Hydrachnidia: Unionicolidae). Ecologica Montenegrina, 54, 53-56.
Rasaily, R., V.J. Kalkman, O. Katel & C. Dorji (2021). Composition of the dragonfly fauna at different altitudes in Bhutan based on larval samples. IDF-report 160: 1-20.
Rasaily, R., V.J. Kalkman, O. Katel & B. Suberi (2021). The distribution, phenology and altitudinal range of dragonflies and damselflies in Bhutan. IDF-report 160: 21-74.
Smit, H., V. Pešić & M.M. Gurung (2022). The water mite genus Sperchon Kramer, 1877 in Bhutan (Acari: Hydrachnidia: Sperchontidae), with the description of three new species. Acarologia, 62 (3), 754-762. https://doi.org/10.24349/pkqk-sd5d.
Smit, H. & M.M. Gurung (2022). Description of the first species of the water mite genus Aturus Kramer, 1875 from the Himalayan Mountain (Acari: Hydrachnidia: Aturidae). Zootaxa, 5169(5): 494-496. https://zoobank.org/urn:lsid:zoobank.org:pub:5D9363D9-9326-4545- 8DE6.
Van Tol J. (2017). Mission report - Entomological fieldwork Bhutan May-June 2017. Naturalis, Leiden.